Three alleged organisers of the March suicide bombings on the Moscow metro have been killed after resisting arrest, Russian officials say.

It was not immediately clear when or where the killings happened. All those involved in the bombings have now been identified, officials said.

Russian leaders previously warned that the masterminds of the attacks, which killed 40 people, would be “destroyed”.

Two young women from Dagestan were identified as carrying out the attacks.

The 29 March bombings targeted two of the Moscow metro’s commuter trains.